Now that the 4Runner is restricted to a V-6 engine, and no V-8 is offered, the fuel economy is somewhat respectable on all models, especially considering the capability the off-road hardware can allow.
The EPA rates rear-wheel-drive 4Runners at 17 mpg city, 22 highway. And with either four-wheel-drive system, the 4Runner returns 17 mpg in the city and 21 mpg on the highway.
There are seven-passenger crossovers that match or exceed those numbers, though. The 4Runner's body-on-frame layout has some advantages off-road or when hauling heavy loads, but its added weight (compared to modern car-based crossovers) and boxier body altogether just aren't as good for gas mileage. In choosing an off-road-oriented vehicle such as this, this is a sacrifice that needs to be taken into consideration.